Page 13 - 《华中农业大学学报（自然科学版）》2022年第5期

第 5 期 王新超 等：中国茶树遗传育种发展、创新之回顾与展望 7

map construction and QTL mapping for flavonoid-related traits 系［J］. 食品科学，2017，38（16）：104-109.DAI W D，XIE D C，
in a tea plant（Camellia sinensis）using 2b-RAD sequencing［J/ LÜ M L，et al.Relationship of flavonol glycoside and processing
OL］.BMC genomics，2018，19：955［2022-02-11］.https：//doi. suitability of tea varieties［J］.Food Sci，2017，38（16）：104-109
org/10.1186/s12864-018-5291-8. （in Chinese）.
［13］ MA J Q，JIN J Q，YAO M Z，et al.Quantitative trait loci map⁃ ［26］ MU B，ZHU Y，LV H P，et al.The enantiomeric distributions of
ping for theobromine and caffeine contents in tea plant（Camellia volatile constituents in different tea cultivars［J］. Food Chem，
sinensis）［J］.J Agr Food Chem，2018，66（50）：13321-13327. 2018，265：329-336.
［14］ HUANG R，WANG J Y，YAO M Z，et al.Quantitative trait loci ［27］ 郭吉春，叶乃兴，何孝延 .茶树杂交一代展叶期的遗传变异［J］.
mapping for free amino acid content using an albino population 茶叶科学，2004，24（4）：255-259.GUO J C，YE N X，HE X Y.
and SNP markers provides insight into the genetic improvement Genetic variation in the leaf-expansion period of the first hybrid
of tea plants ［J/OL］. Hortic Res-England，2022，9：uhab029 generation tea plants.［J］.J Tea Sci，2004，24（4）：255-259（in
［2022-02-11］.https：//doi.org/10.1093/hr/uhab029. Chinese）.
［15］ JIN J Q，YAO M Z，MA C L，et al.Association mapping of caf⁃ ［28］ ZHENG Y，WANG P，CHEN X，et al.Transcriptome and me⁃
feine content with TCS1 in tea plant and its related species［J］. tabolite profiling reveal novel insights into volatile heterosis in
Plant Physiol Bioch，2016，105：251-259. the tea plant（Camellia Sinensis）［J/OL］.Molecules，2019，24：
［16］ JIN J Q，LIU Y F，MA C L，et al.A novel F3'5'H allele with 14 3380 ［2022-02-11］. https：//doi. org/10.3390/mole⁃
bp deletion is associated with high catechin index trait of wild tea cules24183380.
plants and has potential use in enhancing tea quality［J］. J Agr ［29］ XIA E H，ZHANG H B，SHENG J，et al.The tea tree genome
Food Chem，2018，66（40）：10470-10478. provides insights into tea flavor and independent evolution of caf⁃
［17］ WEI K，WANG X，HAO X，et al. Development of a genome- feine biosynthesis［J］.Mol Plant，2017，10（6）：866-877.
wide 200 K SNP array and its application for high-density genetic
［30］ WEI C，YANG H，WANG S，et al. Draft genome sequence of
mapping and origin analysis of Camellia sinensis［J］.Plant Bio⁃
Camellia sinensis var.sinensis provides insights into the evolution
technol J，2022，20：414-416. of the tea genome and tea quality［J］. PNAS，2018，115（18）：
［18］ JIN J Q，MA J Q，YAO M Z，et al.Functional natural allelic vari⁃
E4151-E4158.
ants of flavonoid 3'，5'-hydroxylase gene governing catechin traits
［31］ XIA E，TONG W，HOU Y，et al.The reference genome of tea
in tea plant and its relatives［J］.Planta，2017，245（3）：523-538.
plant and resequencing of 81 diverse accessions provide insights
［19］ WANG R J，GAO X F，YANG J，et al.Genome-wide associa⁃
into its genome evolution and adaptation［J］.Mol Plant，2020，13
tion study to identify favorable SNP allelic variations and candi⁃
（7）：1013-1026.
date genes that control the timing of spring bud flush of tea（Ca⁃
［32］ ZHANG Q J，LI W，LI K，et al.The chromosome-level reference
mellia sinensis）using SLAF-seq［J］.J Agr Food Chem，2019，67
genome of tea tree unveils recent bursts of non-autonomous LTR
（37）：10380-10391.
retrotransposons in driving genome size evolution［J］.Mol Plant，
［20］ FANG K，XIA Z，LI H，et al.Genome-wide association analysis
2020，13（7）：935-938.
identified molecular markers associated with important tea flavor-
［33］ ZHANG W，ZHANG Y，QIU H，et al.Genome assembly of wild
related metabolites ［J/OL］. Hortic Res-England，2021，8：42
tea tree DASZ reveals pedigree and selection history of tea variet⁃
［2022-02-11］.https：//doi.org/10.1038/s41438-021-00477-3.
ies［J/OL］.Nat Commun，2020，11：3719［2022-02-11］.https：//
［21］ LÜ Q，CHEN C，XU Y，et al.Optimization of agrobacterium tu⁃
mefaciens -mediated transformation systems in tea plant（Camel⁃ doi.org/10.1038/s41467-020-17498-6.
［34］ WANG X，FENG H，CHANG Y，et al.Population sequencing
lia sinensis）［J］.Hortic Plant J，2017，3（3）：105-109.
enhances understanding of tea plant evolution［J/OL］.Nat Com⁃
［22］ 唐雨薇，刘丽萍，王若娴，等 . 茶树咖啡碱合成酶 CRISPR/Cas9
mun，2020，11：4447 ［2022-02-11］. https：//doi. org/10.1038/
基因组编辑载体的构建［J］. 茶叶科学，2016，36（4）：414-426.
s41467-020-18228-8.
TANG Y W，LI L P，WANG R X，et al. Development of a
［35］ ZHANG X，CHEN S，SHI L，et al.Haplotype-resolved genome
CRISPR/Cas9 constructed for genome editing of caffeine syn⁃
assembly provides insights into evolutionary history of the tea
thase in Camellia sinensis［J］. Journal of tea science，2016，36
plant Camellia sinensis［J］.Nat Genet，2021，53（8）：1250-1259.
（4）：414-426（in Chinese）.
［23］ 杨亚军 . 茶树育种品质早期化学鉴定Ⅱ. 鲜叶的主要生化组分 ［36］ ZHANG W，LUO C，SCOSSA F，et al.A phased genome based
与 绿 茶 品 质 的 关 系［J］. 茶 叶 科 学 ，1991，11（2）：127-131. on single sperm sequencing reveals crossover pattern and com⁃
YANG Y J.Chemical evaluation on tea quality during early-stage plex relatedness in tea plants［J］.Plant J，2021，105（1）：197-208.
of breeding program Ⅱ . Relationship between the biochemical ［37］ WANG P，YU J，JIN S，et al.Genetic basis of high aroma and
component content in the shoots and the quality of green tea［J］. stress tolerance in the oolong tea cultivar genome［J/OL］.Hortic
J Tea Sci，1991，11（2）：127-131（in Chinese）. Res-England，2021，8：107 ［2022-02-11］. https：//doi. org/
［24］ 杨亚军 . 茶树育种品质早期化学鉴定Ⅰ. 鲜叶的主要生化组分 10.1038/s41438-021-00542-x.
与红茶品质的关系［J］. 茶叶科学，1990，10（2）：59-64.YANG ［38］ XIA E H，TONG W，WU Q，et al.Tea plant genomics：achieve⁃
Y J. Chemical evaluation on tea quality during early-stage of ments，challenges and perspectives［J/OL］.Hortic Res-England，
breeding program Ⅰ.Relationship between the biochemical com⁃ 2020，7：7［2022-02-11］. https：//doi. org/10.1038/s41438-019-
ponent content in the shoots and the quality of black tea［J］. J 0225-4.
Tea Sci，1990，10（2）：59-64（in Chinese）. ［39］ WANG L，YAO L N，HAO X Y，et al.Transcriptional and phys⁃
［25］ 戴伟东，解东超，吕美玲，等 . 黄酮醇糖苷与茶树品种适制性关 iological analyses reveal the association of ROS metabolism with